• Phylogenetic placement: Passeri: Passerides: Sylviida: Aegithaloidea
• Distribution: Africa, Eurasia, Wallacea, North America
• Number of extant genera: 1
• Number of extant species: 80

Traditional genus-level classification of extant Phylloscopidae following the AviList checklist v2025. (link) The number of subspecies is given in parentheses.  

[Notes: (1) It is unclear whether P. emilsalimi and P. suaramerdu, erected by Rheindt et al., 2020, should be included. (2) an additional species, P. tokaraensis, was described by Saitoh et al., 2026 ]

References

Alström P, Ericson PGP, Olsson U, and Sundberg P (2006), Phylogeny and classification of the avian superfamily Sylvioidea, Mol. Phylogenet. Evol. 38, 381-397. (abstract)

Alström P, Rheindt FE, Zhang R, Zhao M, Wang J, Zhu X, Gwee CY, Hao Y, Ohlson J, Jia C, Prawiradilaga DM, Ericson PGP, Lei F, and Olsson U (2018), Complete species-level phylogeny of the leaf warbler (Aves: Phylloscopidae) radiation, Mol. Phylogenet. Evol. 126, 141-152. (abstract) [note: study based on one mitochondrial and three nuclear loci]

Barker FK, Barrowclough GF, and Groth JG (2002) A phylogenetic hypothesis for passerine birds: taxonomic and biogeographic implications of an analysis of nuclear DNA sequence data, Proc. R. Soc. B 269, 295-308 (abstract)

Barker FK, Cibois A, Schikler P, Feinstein J, and Cracraft J (2004) Phylogeny and diversification of the largest avian radiation, Proc. Natl. Acad. Sci. U.S.A. 101, 11040-45. (pdf)

DeCicco LH, DeRaad DA, Holland P, Pikacha D, Tigulu IG, Tako R, Boseto D, Sweet PR, and Miyle RG (2024), Phylogeography of the Island Leaf Warbler (Aves: Phylloscopus poliocephalus) in northern Melanesia reveals rapid secondary sympatry or ecological speciation on Kolombangara Island, Solomon Islands, Bull. Soc. Syst. Biol. 3. (pdf)

Fregin S, Haase M, Olsson U, and Alström P (2012), New insights into family relationships within the avian superfamily Sylvioidea (Passeriformes) based on seven molecular markers, BMC Evol. Biol. 12, e:157. (free pdf)

Gräf L, Griebeler EM, Oldeland J, and Tietze DT (2025), Climate-niche evolution in leaf-warblers (Aves: Phylloscopidae): a matter of phylogeny, Diversity 17, e:844. (free pdf) [note: compares the phylogenies of Tietze et al. 2015 and Alström et al. 2018]

Johansson US, Fjeldså J, and Bowie RCK (2008), Phylogenetic relationships within Passerida (Aves: Passeriformes): A review and a new molecular phylogeny based on three nuclear intron markers, Mol. Phylogenet. Evol. 48, 858-876. (abstract)

Ng NSR, Prawiradilaga DM, Ng EYX, Suparno, Ashari H, Trainor C, Verbelen P, and Rheindt FE (2018), A striking new species of leaf warbler from the Lesser Sundas as uncovered through morphology and genomics, Sci. Rep. 8, e:15646. (free pdf)

Olsson U, Alström P, Ericson PGP, and Sundberg P (2005), Non-monophyletic taxa and cryptic species - evidence from a molecular phylogeny of leaf-warblers (Phylloscopus, Aves), Mol. Phylogenet. Evol. 36, 261-276. (abstract)

Päckert M, Martens J, and Sun YH (2010), Phylogeny of long-tailed tits and allies inferred from mitochondrial and nuclear markers (Aves: Passeriformes, Aegithalidae), Mol. Phylogenet. Evol. 55, 952-967. (abstract)

Rheindt FE, Prawiradilaga DM, Ashari H, Suparno, Gwee CY, Lee GWX, Wu MY, and Ng NSR (2020), A lost world in Wallacea: description of a montane archipelagic avifauna, Science 367, 167-170. (pdf) [note: paper includes description of Phylloscopus suaramerdu and P. emilsalimi]

Saitoh T, Shipilina D, Xia C, Zhang L, Seki SI, Olsson U, and Alström P (2026), Discovering and protecting cryptic biodiversity: A case study of a previously undescribed, vulnerable bird species in Japan, PNAS Nexus 5, e:pgag037. (free pdf) [note: described a new species, Phylloscopus tokaraensis, split from P. ijimae; and provided a mitochondrial cytochrome b tree for all Phylloscopus species in S7]

Tietze DT, Martens J, Fischer BS, Sun Y, Klussmann-Kolb A, and Päckert M (2015), Evolution of leaf-warbler songs (Aves: Phylloscopidae), Ecol. Evol. 5, 781-798. (pdf)