The Passeriformes, or perching birds represent almost two-thirds of all bird species. For this reason, this order is only treated to family-level. Nevertheless, the Passeriformes are presented here in two timetrees, with the Passerida being treated in a separately.


Timetree of Passeriformes based on Oliveros et al. (2019) and Kuhl et al. (2021), with the distribution of each taxon being indicated by the colour-code used throughout this website (see Distribution code). In contrast to their treament, however, Lamproliidae are treated here as a separate family rather than a subfamily of Rhipiduridae. Passeri and Tyranni together constitite the Eupasseres. The few differences between the Oliveros et al. (2019) and the Kuhl et al. (2021) trees are indicated by dashed lines. Divergence times follow Kuhl et al. (2021). 


The phylogenetic interrelations among Passerides are outlined in the timetree below:

Timetree of Passerides based on Oliveros et al. (2019), and Kuhl et al., (2021), with the distribution of each taxon being indicated by the colour-code used throughout this website (see Distribution code). The phylogenetic relationships among Sylvioidea have been further detailed by Cai et al. (2019). In contrast to Oliveros et al. (2019), a number of taxa have been upgraded to family status in the above cladogram: Modulatricidae, Cettiidae, Erythrocercidae, Chaetopidae, and Picathartidae. The few differences between the Oliveros et al. (2019) and the Kuhl et al. (2021) trees are indicated by dashed lines. Divergence times follow Kuhl et al. (2021). 


Alström, P., D.M. Hooper, Y. Liu, U. Olsson, D. Mohan, M. Gelang, H.L. Manh, J. Zhao, F. Lei, and T.D. Price (2014), Discovery of a relict lineage and

monotypic family of passerine birds, Biol. Lett. 10, 20131067. DOI: 10.1098/rsbl.2013.1067. (pdf)

Alström, P., F.E. Rheindt, R. Zhang, M. Zhao, J. Wang, X. Zhu, C.Y. Gwee, Y. Hao, J. Ohlson, C. Jia, D.M. Prawiradilaga, P.G.P. Ericson, F. Lei, and U.

Olsson. Price (2018), Complete species-level phylogeny of the leaf warbler (Aves: Phylloscopidae) radiation, Mol. Phylogenet. Evol. 126, 141-152. DOI: 10.1016/j.ympev.2018.03.031. (abstract)

Barker, F.K. (2014), Mitogenomic data resolve basal relationships among passeriform and passeridan birds, Mol. Phylogenet. Evol. 79, 313-324. DOI:

10.1016/j.ympev.2014.06.011. (abstract)

Barker, F.K. (2017), Molecular phylogenetics of the wrens and allies (Passeriformes: Certhioidea), with comments on the relationships of Ferminia, Amer.

Mus. Novitates 3887, 1-28. (pdf)

Cai, T., A. Cibois, P. Alström, R.G. Moyle, J.D. Kennedy, S. Shao, R. Zhang, M. Irestedt, G.P. Ericson, M. Gelang, Y. Qu, F. Lei, and J. Fjelda (2019), Near-

complete phylogeny and taxonomic revision of the world´s babbler (Aves: Passeriformes), Mol. Phylogenet. Evol. 130, 346-356. DOI: 10.1016/j.ympev.2018.10.010. (abstract)

Harvey, M.G., G.A. Bravo, S. Claramunt, A.M. Cuervo, G.E. Derryberry, J. Battilana, G.F. Seeholzer, J.S. McKay, B.C. O'Meara, B.C. Faircloth, S.V.

Edwards, J. Perez-Eman, R.G. Moyle, F.H. Sheldon, A. Aleixo, B.T. Smith, R.T. Chesser, L.F. Silveira, J. Cracraft, R.T. Brumfield, and E.P. Derryberry (2020), The evolution of a tropical biodiversity hotspotScience 370, 1343-1348. DOI: 10.1126/science.aaz6970. (abstract)

Jønsson, K.A., P.-H. Fabre, J.D. Kennedy, B.G. Holt, M.K. Borregaard, C. Rahbek, and J. Fjeldså (2016), A supermatrix phylogeny of corvoid passerine

birds (Aves: Corvides), Mol. Phylogenet. Evol. 94, 87-94. DOI: 10.1016/j.ympev.2015.08.020. (abstract)

Kuhl, H., C. Frankl-Vilches, A. Bakker, G. Mayr, G. Nikolaus, S.T. Boerno, S. Klages, B. Timmermann, and M. Gahr (2021), An unbiased molecular approach

using 3'UTRs resolves the avian family-level tree of life, Mol. Biol. Evol. 38(1), 108-127. DOI: 10.1093/molbev/msaa191. (pdf)

Mackiewicz, P., A.D. Urantówka, A. Kroczak, and D. Mackiewicz (2019), Resolving phylogenetic relationships within Passeriformes based on mitochondrial

genes and inferring the evolution of their mitogenomes in terms of duplications, Genome Biol. Evol. 11(10), 2824-2849. DOI: 10.1093/gbe/evz209. (pdf)

Marki, P.Z., K.A. Jonsson, M. Irestedt, J.M.T. Nguyen, C. Rahbek, and J. Fjeldsa (2017) Supermatrix phylogeny and biogeography of the Australasian

Meliphagides radiation (Aves: Passeriformes), Mol. Phylogenet. Evol. 107, 516-529. DOI: 10.1016/j.ympev.2016.12.021. (abstract)

Moyle, R.G., C.H. Oliveros, M.J. Andersen, P.A. Hosner, B.W. Benz, J.D. Manthey, S.L. Travers, R.M. Brown, and B.C. Faircloth (2016), Tectonic collision

and uplift of Wallacea triggered the global songbird radiation, Nature Comm. 7, 12709. DOI: 10.1038/ncomms12709. (pdf)

Oliveros, C.H., D.J. Field, D.T. Ksepka, K.F. Barker, A. Aleixo, M.J. Andersen, P. Alström, B.W. Benz, E.L. Braun, M.J. Braun, G.A. Bravo, R.T. Brumfield,

R.T. Chesser, S. Claramunt, J. Cracraft, A.M. Cuervo, E.P. Derryberry, T.C. Glenn, M.G. Harvey, P.A. Hosner, L. Joseph, R.T. Kimball, A.L. Mack, C.M. Miskelly, A.T. Peterson, M.B. Robbins, F.H. Sheldon, L.F. Silveira, B.T. Smith, N.D. White, R.G. Moyle, and B.C. Faircloth (2019), Earth history and the passerine superradiation, PNAS 116 (16), 7915-7925. DOI: 10.1073/pnas.1813206116. (pdf)

Selvatti, A.P., L.P. Gonzaga, C.A. de Moraes Russo (2015), A Paleogene origin for crown passerines and the diversification of the Oscines in the New World,

Mol. Phylogenet. Evol. 88, 1-15. DOI: 10.1016/j.ympev.2015.03.018. (pdf)

Selvatti, A.P., A. Galvão, A.G. Pereira, L.P. Gonzaga, and C.A. de Moraes Russo (2017), An African origin of the Eurylaimides (Passeriformes) and the

successful diversification of the ground-foraging pittas (Pittidae), Mol. Biol. Evol. 34(2), 483-499. DOI: 10.1093/molbev/msw250. (pdf)

Slager, D.L., C.J. Battey, R.W. Bryson, G. Voelker, and J. Klicka (2014), A multilocus phylogeny of a major New World avian radiation: the Vireonidae, Mol.

Biol. Evol. 80, 95-104. DOI: 10.1016/j.ympev.2014.07.021. (abstract)

Suh, A., S. Bachg, S. Donnellan, L. Joseph, J. Brosius, J.O. Kriegs and J. Schmitz (2017), De-novo emergence of SINE retroposons during the early evolution

of passerine birds, Mobile DNA 8:21. DOI:10.1186/s13100-017-0104-1.(pdf)


Blue-faced Honeyeater (Entomyzon cyanotis), Meliphagidae (Foto: Carolin Pfeiffer)

Montezuma Oropendula (Psarocolius montezuma), Icteridae

White-eared Catbird (Ailuroedus buccoides), Ptilonorhynchidae (Foto: Carolin Pfeiffer)

Hooded Pitta (Pitta sordida), Pittidae (Foto: Carolin Pfeiffer)

Andean Cock-of-the-rock (Rupicola peruviana), male, Cotingidae (Foto: Carolin Pfeiffer)

Gouldian Finch (Erythrura gouldiae), Estrildidae (Foto: Carolin Pfeiffer)

Vermilion Flycatcher (Pyrocephalus rubinus), Tyrannidae (Foto: Carolin Pfeiffer)

Chestnut-backed Thrush (Geokichla dohertyi), Turdidae (Foto: Carolin Pfeiffer)

Black-and-white Mannikin (Lonchura bicolor), Estrildidae (Foto: Carolin Pfeiffer)

Andean Cock-of-the-rock (Rupicola peruviana), female, Cotingidae (Foto: Carolin Pfeiffer)

Brazilian Tanager (Ramphocelus bresilius), Thraupidae (Foto: Carolin Pfeiffer)

Paradise Tanager (Tangara chilensis), Thraupidae (Foto: Carolin Pfeiffer)